What is a PCIe slot with an x4 slot and x16 connector?

pcie slot x16
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Physically yes, the slot connector for an x4 is generally smaller and PCIe is designed that smaller cards can occupy larger slots if needed

Have you ever thought if you can plug PCI Express cards into x16 lane slot? In this video

pcie 4 in pcie 3 slot Buy PCI Express X1 to X16 Low Profile Slot Extension Adapter

pcie x1 slot function PCIe x16 Gen expansion backplane with two slots The PCIe x16 Gen3 backplane has two PCI Express x16 slots, with one slot supporting the OSS target